Ando K., Yamasako J. 2013. Study of Tenebrionid Fauna of Sulawesi. I. Tribes Bolitophagini, Trachyscelini, and Subtribe Heterocheirina of the Opatrini (Coleoptera, Tenebrionidae). Elytra. New Series. 3: 275–294.

Ando K., Yamashita S., Meleng P., Itioka T. 2022. Tenebrionidae (Coleoptera) collected from fruiting bodies of polypores in Sarawak, Malaysia. Contributions from the Biological Laboratory, Kyoto University. 31: 159–210.

Bouchard P., Bousquet Y., Aalbu R. L., Alonso-Zarazaga M. A., Merkl O., Davies A. E. 2021. Review of genusgroup names in the family Tenebrionidae (Insecta, Coleoptera). ZooKeys. 1050: 1–633.

Gebien H. 1914. I. - Die Tenebrioni-denfauna Borneos.—Erster Teil. The Sarawak Museum Journal. 2, part 1: 1–58, pl. 1.

Gebien H. 1922. Philippine Tenebrionidae, II. The Philippine Journal of Science. 19: 439–515.

Gebien H. 1925 Die Tenebrioniden (Coleoptera) des indomalayischen Gebietes, unter Beruecksichtigung der benachbarten Faunen, II. Die Gattungen Atasthalus, Bolitoxenus, Bolitonaeus, und Sumbawia. The Philippine Journal of Science. 26: 423–44, pls. 1–2.

Gebien H. 1944. Katalog der Tenebrioniden. Teil III. Mitteilungen der Münchener Entomologischen Gesellschaft. 34: 497–554 (842–899).

Grimm R. 2014. New and little known species of Tenebrionidae (Coleoptera) from Borneo (4). Stuttgarter Beiträge zur Naturkunde A. Neue Serie. 7: 183–197.

Iwan D., Löbl I., Bouchard P., Bousquet Y., Kamiński M., Merkl O., Ando K., Schawaller W. 2020. Family Tenebrionidae Latreille, 1802: 104–475. In: Iwan D., Löbl I. (eds.): Catalogue of Palaearctic Coleoptera. Volume 5. Revised and Updated Second Edition. Tenebrionoidea. Brill, Leiden & Boston. Pp. xxiv + 945.

Jung H.B. 2012. Taxonomic Review of the Fungivorous Tenebrionid Boletoxenus Motschulsky, 1858 (Coleoptera: Tenebrionidae: Tenebrioninae: Bolitophagini) in Korea with Host Fungi. Korean Journal of Applied Entomology. 51: 265–269.

Kaszab Z. 1965. Wissenschaftliche Ergebnisse der von Dr. F. Schmid in Indien gesammelten Tenebrioniden (Coleoptera). Miscelánea Zoológica. 2: 107–130.

Masumoto K. 1982. Tenebrionidae of Formosa (4). Elytra. 10: 17–32.

Masumoto K., Akita K. 2001. Two new tenebrionid species from Japan. Special Publication of the Japan Coleopterological Society. 1: 247–250.

Masumoto K., Akita K. 2003. New or littleknown tenebrionid species from Japan (Part 4) (Coleoptera). Entomological Review of Japan. 58: 173–182.

Masumoto K., Akita K., Lee C.-F. 2008: New tenebrionid beetles from Taiwan (4). Entomological Review of Japan. 63: 15–26.

Motschulsky V. de 1858. Entomologie spéciale. Insectes des Indes orientales. 1:ière Série. Études Entomologiques. 7: 20–122, 2 pls.